PifferPilfer

PifferPilfer

Were Ancient Elites Genetically Different or Just Related?

Burial richness after close-kin pruning and comparisons between and within families

Davide Piffer's avatar
Davide Piffer
Oct 30, 2026
∙ Paid
AI-generated illustration of richly and sparsely furnished ancient graves linked by branching golden lines symbolising family connections.
Illustration of unequal grave furnishing and shared family connections. The scene is imagined and does not reconstruct a particular cemetery.

A richly furnished grave makes me wonder what set its occupant apart. In my earlier post, Digging the elite: richer graves, lighter skin?, I explored whether people buried with more goods also differed genetically. Richer burials were associated with higher fluid-intelligence scores and a pigmentation score pointing toward lighter untanned skin. These scores summarise genetic variants associated with traits in modern studies; fluid intelligence refers to reasoning through unfamiliar problems.

But a cemetery can preserve a family’s history as much as a community’s social hierarchy. Imagine several relatives receiving elaborate burials. They share DNA and may also share inherited standing. Counting them as independent examples could make one dynasty look like a general connection between genes and privilege.

Since that post, I have checked additional burial inventories against their sources and combined 406 newly included people with the corrected earlier records. The resulting dataset contains 1,366 people. I first revisit the earlier associations, including the score linked to education today, before testing how much they depend on relatives.

I approach the family question in three ways. First, I remove close relatives until no documented first- or second-degree pair remains and repeat the analysis. Then I compare relatives buried differently in the same cemetery, with a separate check for brothers and sisters. Finally, I separate differences between family averages from differences among relatives in one model.


A family behind the comparisons

Figure 1 shows a small branch of an Avar-period family at Rákóczifalva in Hungary. The father, RKC011, the mother, RKC013, and their daughter, RKC029, all have genetic records and burial scores in this analysis. Their graves received different furnishing. The links come from the source-supported pedigree in the dataset, checked against Gnecchi-Ruscone et al. (2024).

Figure 1. Parents, their daughter and different burial scores

A source-supported Rákóczifalva pedigree branch shows father RKC011, mother RKC013 and daughter RKC029, with burial object-value scores zero, two and one.
The diagram joins the documented father–daughter and mother–daughter links to each person’s contextual object-value index (COVI). The daughter was a juvenile at death; both parents were adults. Only this branch of the larger pedigree is shown. The values measure recorded burial furnishing, rather than object counts or lifetime wealth. This illustrates the family comparison; the reported associations use the full eligible family sample. Source: Gnecchi-Ruscone et al. (2024), Supplementary Figure 10, matched to the analysis dataset.

Between-group models compare average genetic scores with average burial treatment across documented kin groups in the same cemetery. Everyone eligible stays in the comparison, including a group represented by one person. Within-family models compare relatives against their own family’s baseline: does the person with the higher genetic score receive the richer burial? The kin groups follow documented biological relationships from ancient-DNA studies and verified pedigrees; shared burial style alone does not define a family.

The initial cemetery models contain both kinds of difference. The family models focus on differences among relatives buried in the same cemetery. I then separate the between-family and within-family components explicitly. The logic of the comparison among relatives resembles a within-family genome-wide association study (GWAS): relatives inherit different combinations of variants. Here I test existing genetic scores against burial treatment, rather than scanning individual variants across the genome.

Figure 2 compares the earlier records, the enlarged dataset, two ways of removing close relatives, and the comparison within families. The earlier measurement scales are retained throughout.

Figure 2. Burial furnishing and genetic scores before and after the family checks

Forest plot comparing earlier and expanded burial cohorts, kin-pruned samples and within-family estimates for education, fluid intelligence and lighter-skin scores.
Outcome: log(1 + object-value index). Filled circles show ordinary regression with cemetery-clustered standard errors; hollow squares show a mixed model fitted with a genetic relationship matrix (GRM). The latter models genetic similarity between individuals and can change both coefficients and intervals. Both retain the same people and covariates. The family estimates compare relatives buried differently in the same cemetery.

Figure 2 and the tables use the earlier within-cemetery standard-deviation units, with 95% confidence intervals.

Does inherited family standing explain the whole pattern, or do differences between relatives still show through? The fluid-intelligence result gives me a particularly interesting lead. Pigmentation brings a large Avar family into focus for a different reason. Following those two clues gets us closer to the question behind the rich graves: what came with the family, and what distinguished the individual?

Upgrade to see what survives when close relatives are removed—and why fluid intelligence and pigmentation tell different stories inside ancient families.

User's avatar

Continue reading this post for free, courtesy of Davide Piffer.

Or purchase a paid subscription.
© 2026 Davide Piffer · Privacy ∙ Terms ∙ Collection notice
Start your SubstackGet the app
Substack is the home for great culture